Episode 4 of Season 2 of "Mayfair Witches" kicks off with Rowan immersed in a dream that revisits her tumultuous past with Lark, revealing for the first time that their breakup occurred years prior due to Lark's unilateral decision to apply for a job in New York without consulting her.
Moira interrupts Rowan's slumber, urgency lacing her voice as she tasks Rowan with resolving the tricky situation between Jojo and Daphne. Rowan attempts to wield her powers, but without Lasher by her side, she finds herself woefully inadequate to extricate the pair from their enchantment. As they ponder their next steps, Lark's call pierces through the tension. He expresses concern over distressing messages Rowan had sent the previous night and reminded her of their impending meeting before his departure. Rowan, sensing the gravity of the situation, enlists Moira to contact Ciprien and bring Lasher within proximity, hoping to harness his power to rectify the Jojo-Daphne predicament.
However, fate conspires against them; Albrecht, with Lasher in tow, complicates matters further. Using his psychic abilities, Ciprien discovers Albrecht's treacherous betrayal of The Talamasca and his subterfuge in spiriting Lasher away. Understandably, Ciprien's fury knows no bounds upon realizing he has once again been manipulated by Albrecht.
Meanwhile, Rowan ventures out to meet Lark, their conversation a nostalgic ballet of shared memories until Lark gently broaches the subject of Rowan's recent erratic behavior. She confides in him, her voice tinged with anxiety, about a string of mysterious deaths plaguing her family, though she remains vague on the specifics. Rowan expresses gratitude for his support, and Lark vows to stand by her side, determined not to let her down again.
Back at the grand Mayfair mansion, Cortland remains confined within his father's room, haunted by Millie's ghostly presence. Moira, left to grapple with the ongoing crisis, strives to reach Ciprien while seeking a means to communicate with Daphne. Her efforts are momentarily sidetracked by a call from Ryan, a family member extending an invitation to a gathering fraught with tension. The household is in uproar, each member wary of Rowan and intent on safeguarding themselves. As their conversation unfolds, the mansion begins to crumble around them, prompting Moira to urgently summon Rowan. Rowan, reluctantly bidding farewell to Lark, rushes back to the crumbling estate.
Unbeknownst to Rowan, Lark retains her cup, using it as a pretext to gain access to a nearby laboratory through his office. Upon her return, Rowan is astounded by the mansion's dilapidated state. In unison, she and Moira decide to confront Cortland, hoping to unlock the secrets he holds about Lasher. Employing her telepathic abilities, Moira uncovers that Cortland's collaboration with Albrecht stems from his loyalty to his father, Julian. Yet, despite her probing, Cortland remains in the dark about Lasher's true intentions. Intriguingly, Moira observes a palpable fear of his father lurking in Cortland's eyes.
As Cortland hastily attempts to drown out Moira's haunting thoughts with the soothing embrace of music, they both come to an abrupt realization: Julian's vintage Victrola, a piece steeped in nostalgia, is conspicuously absent. Their minds swiftly converge on Dolly Jean, who had spirited it away under the cover of darkness the previous evening. Rowan, sensing the urgency, urges Cortland to remain and devise a contingency plan, should Dolly Jean prove unforthcoming with answers.
Meanwhile, within the hallowed halls of the Talamasca, Ciprien stumbles upon Albrecht's final missive, tucked away in his office. Its contents chill him to the bone – Albrecht had harbored treacherous designs against their very own organization. Determined to uncover the motive, Ciprien delves deeper, only to discover a charred scrap of paper in the fireplace, barely escaping the flames' clutches. But before he can decipher its cryptic message, Polina, a new agent with an air of suspicion, enters the scene. She grills Ciprien relentlessly, her conviction unshakeable that he was in league with Albrecht, and escorts him for further interrogation by the elders.
Elsewhere, Rowan and Moira find themselves at Dolly Jean's abode, where she sits in a trance-like state, her spirit wandering elsewhere. Evelyn, sensing their intent to snap her out of it, intervenes gently, explaining that Julian, before his untimely demise, had forged an unbreakable bond with the antique Victrola. Through it, Dolly Jean and Evelyn take turns to nurture their relationship with Julian beyond the veil of mortality. Time passes in a blur, as Dolly Jean and Julian's ethereal encounters unfold, until her allotted time draws to a close. Evelyn, with tender care, gently pulls Dolly Jean back into the realm of the living.
Safely reunited with Dolly Jean, Rowan interrogates her about Julian's potential insights into Lasher, hinting that he might hold the key to understanding the latter's intentions. Dolly Jean confides that Julian is indeed the one individual who could shed light on Lasher's motives. Nevertheless, Julian insists on his ignorance since Albrecht's departure with Lasher. Rowan and Moira remain skeptical of Julian's claims. Determined to confront Julian, Rowan insists on entering the mystical antique victrola. Dolly Jean and Evelyn attempt to dissuade her, warning of Julian's potent wizardry even in death, a force unforgiving towards its adversaries.
Amidst their heated exchange, Moira abruptly dashes towards the victrola and slips into a trance. Her cries escalate, prompting Rowan to wrench her away despite the sisters' protests. Moira struggles to breathe, necessitating an urgent trip to the hospital, as Rowan, devoid of healing abilities, watches helplessly.
Meanwhile, back at the Mayfair mansion, Millie persists in tormenting Cortland, still wounds raw from her father's betrayal of her beloved. She lays the blame on Cortland for mirroring their father's nature. Yet, Cortland manages to convince Millie of his genuine affection for Jojo and his desire to aid her. He persuades Millie to relay a message to Jojo, and she, albeit reluctantly, agrees to assist him.
Once Moira regains consciousness, Rowan confides in her about her predicament with Lasher. Moira comes to empathize with Rowan, their shared Mayfair traumas forging a bond between them. Moira volunteers to be Rowan's anchor, and they, accompanied by Cortland, venture back to Dolly Jean's abode.
Elsewhere, Lark receives the results of his latest test, revealing an intriguing connection between Rowan and the sample she provided.
Returning to Dolly Jean's house, Rowan steps into the antique victrola to confront Julian. Warnings echo around her to tread carefully, and the episode concludes on a tense note as Julian grants her entry.